On the ball
Meaning: To be alert; in command of one's senses.
Origin: Some authorities have suggested that 'on the ball' originated in the sporting arena, and alludes to runners being on the balls of their feet, eagerly ready to run a race. This has some similarities with being 'up to scratch', which derives from boxers or runners being ready at the starting line. It is a plausible derivation, but has nothing to recommend it beyond that.
Need I go on? It isn't true.
The phrase 'on the ball' did actually originate in the sporting arena, but relates to the eyes rather than the feet. It is a contraction of the earlier expression 'keep your eye on the ball', which advice has been given to participants in virtually every known ball game. For the source, we need to look to early ball games. The phrase is recorded in early records of cricket, golf, croquet and baseball and many people regard baseball as the origin. Well, that appears to be almost true - the earliest citation that I can find in print comes from the English game of rounders. The English novelist William Kingston wrote 'books for boys', and in 1864 published Ernest Bracebridge, or, Schoolboy Days, which includes this scene:
Ellis seized the bat with a convulsive clutch... Remembering Ernest's advice, he kept his eye on the ball, and hit it so fairly that he sent it flying away to a considerable distance. "Capital!" cried Ernest. "Run! run! - two bases at least."
American readers will recognise the similarity of the rounders terminology with that of baseball. For those not familiar with rounders and/or baseball, suffice it to say that they are essentially the same game, but that it is easier to imagine Sylvester Stallone playing baseball. There's no consensus on this but there's a strong case to be made that baseball is in fact an English game, being merely a beefed-up variation of rounders.
The ball, once stuck off,
Away flies the boy
To the next destin'd post,
And then home with joy.
Baseball may or may not have been the origin of 'keep your eye on the ball', but it did take over the use of the phrase. As well as as the batters 'keeping their eye on the ball', the pitchers were also said to 'put something on the ball', i.e. they imparted some spin or curve on it. This usage dates from the start of the 20th century, for example, this piece from The Indianapolis Star, April 1910:
Graham put something on the ball that fooled even Bowerman.
The figurative version of the phrase 'on the ball', i.e. with the meaning of being 'alert or apt' in a context where no actual ball is present, began later still. In 1989, W. C. Williams and J. Laughlin published Selected Letters, which contained an extract from a letter written Williams in 1939:
The novella by Quevedo... [is] right on the ball.
As to whether the phrase originated in the USA or the UK, on present evidence, I'd call it a 1-1 draw.
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