The process is explained further at Science.
Infected with a virus that delivers the gene encoding the optogenetic molecular switch, the modified cardiac cells twitch when blue light shines on them. But translating that effect into coherent motion took months of tweaking; simply getting a robot ray to move forward when light stimulated the front of its fin took Park 200 tries. Ultimately, he built 100 more robots and showed they could navigate underwater obstacle courses. To negotiate turns, Park guides a ray with two light sources, one pointed at each fin. Changing the frequency of the light slows or speeds up the contraction rate; by making one side beat faster than the other, he steers the robot left or right.See another video at Popular Mechanics.
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